Lilith, also designated as Punk-02, is one of six satellites of Dr. Vegapunk, embodying his aspect of "evil". Lilith has the appearance of a young woman with wild light hair. She wears a light full-body jumpsuit that bears her designation of Punk-02 on the left breast; she also wears a dark coat, boots, gloves, and aviator helmet. Like the other satellites, she wears an earpiece and antenna on her left ear which enables communication with Vegapunk's other manifestations. Being designated the "evil" satellite of Vegapunk, Lilith is the manifestation of Vegapunk's ruthless and malevolent traits. She utilizes her technology and weaponry to steal from voyagers, chiefly pirates, who trespass on Egghead, justifying this to Shaka as necessary for the sake of keeping Vegapunk's research operational.

However, she possesses a rash attitude which can cause her to underestimate those she attacks, as she was stunned when Shaka, the manifestation of Vegapunk's logic, pointed out that the Straw Hats were equipped and poised to deal great harm to herself and her army of Sea Beasts; in response, Lilith seems capable of heeding Shaka's advice making her far from unreasonable. She uses the first person pronoun washi, which is usually associated with old men in fiction. She also uses stereotypical elderly speech patterns such as the copula ja. Being one of six manifestations of the genius scientist Vegapunk, Lilith has contributed to his numerous technological advances and breakthroughs. Despite the technology and weaponry at her disposal, Lilith was cautioned by Shaka that she was not equipped to take on Roronoa Zoro and Nico Robin, two members of an Emperor's crew with bounties around 1 billion beli, in combat. Lilith has made heavy use of mechas and cybernetically-enhanced animals to aid her in combat. She pilots a giant mecha called the Vega Force 1, which is extremely large to the point of being able to carry the Thousand Sunny in its hands. The Vega Force is able to move underwater and unleash explosions when it punches. Additionally, Lilith has power over Sea Beasts that she has turned into cyborgs. These Sea Beasts are equipped with numerous weapons, with an example being the Mecha-Shark having a torpedo launcher in its mouth. This army of cyborg creatures allows Lilith to easily surround and overwhelm ships sailing the sea, enabling her to loot them. However, she has had issues with achieving programming to completely override the organic urges of the creatures, which limits her ability to have them do exactly what she wants; Shaka himself would note that any equipment, weaponry and modified Sea Beasts at Lilith's disposal would be useless against a swordsman of Zoro's caliber. When the Straw Hat Pirates drew near to Egghead on the Thousand Sunny, the Mecha-Shark went to attack them. With the shark's primal urges overriding its programming and threatening to completely annihilate the crew's possessions, Lilith went underwater in the Vega Force and took it down before lifting the Sunny back above water. Lilith then exited the Vega Force and introduced herself as Dr. Vegapunk, to general confusion. Lilith informed the pirates that she did not rescue them, but rather spared them from the Mecha-Shark so she could rob them of their valuables. She summoned her army of weaponized Sea Beasts to surround the Straw Hats, ordering them to surrender to her their valued possessions at once, but then received a call from fellow satellite Shaka who informed her she had blundered in her antagonism of the pirates, pointing out that they were poised to target and overwhelm her if they retaliate much to Lilith's dismay, embarrassed that she overlooked something that could have easily led to her downfall. Roronoa Zoro told Lilith to play nice, and Shaka told his counterpart to bring the pirates to him as he has interest with the Straw Hats.

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